Moroni, Edson, Department of Biological and Environmental Sciences, Federal University of Grande Dourados – UFGD, Rod. Dourados/Itahum, km 12, 79804‐970, Dourados, MS, Brazil. Department of Hydrobiology, Federal University of São Carlos – UFSCar, Rod. Washington Luís, km 235, 13565‐905, São Carlos, SP, Brazil., Brazil